Many insects have evolved refined self-cleaning membrane structuring to contend with an environment that presents a range of potential contaminates. Contamination has the potential to reduce or interfere with the primary functioning of the wing membrane or affect other wing cuticle properties, (for example, antireflection). Insects will typically encounter a variety of airborne contaminants which include plant matter and soil fragments. Insects with relatively long or large wings may be especially susceptible to fouling due to the high-wing surface area and reduced ability to clean their extremities.
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